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Pleistocene Geology of Kansas

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Stratigraphic Paleontology, continued

Molluscan Faunal Assemblages in Illinoian Deposits

The Crete and Loveland members of the Sanborn formation are the depositional representatives of Illinoian time in Kansas, and since the two members are separable only on a textural basis and are gradational with respect to each other, the molluscan fauna discussed here is a composite assemblage from Crete-Loveland deposits. The occurrence and distribution of 25 kinds of mollusks at 10 localities in the State is shown on Figure 8, and illustrations of a representative assemblage appear on Plate 16.

Figure 8--Occurrence of fossil mollusks in Illinoian deposits at 10 localities in Kansas. An Acrobat PDF version of this figure is available.

Molluscan species Faunal localities
SW sec. 35, T. 17 S., R. 13 W., Barton SW sec. 33, T. 13 S., R. 5 E., Geary NW sec. 21, T. 24 S., R. 21 W., Hodgeman NE sec. 5, T. 2 S., R. 7 W., Jewell NW sec. 2, T. 2 S., R. 9 W., Jewell SE sec. 13, T. 1 S., R. 9 W., Norton NW sec. 23, T. 2 S., R. 18 W., Phillips NW sec. 9, T. 1 S., R. 19 W., Phillips NW sec. 9, T. 2 S., R. 16 W., Phillips NW sec. 7, T. 18 S., R. 7 W., Rice
Anodonta sp.         Circle          
Carychium exiguum (Say)   Circle   Circle     Circle   Circle  
Carychium perexiguum Baker Circle             Circle   Circle
Deroceras laeve (Muller)   Circle         Circle Circle   Circle
Discus cronkhitei (Newcomb)             Circle   Circle  
Gastrocopta armifera (Say)   Circle   Circle     Circle Circle    
Gastrocopta holzingeri (Sterki)   Circle             Circle  
Gastrocopta tappaniana (C. B. Adams)         Circle   Circle      
Gyraulus similaris (Baker)                    
Hawaiia minuscula (Binney)   Circle   Circle       Circle Circle  
Helicodiscus parallelus (Say)   Circle   Circle Circle   Circle      
Helicodiscus singleyanus (Pilsbry)       Circle     Circle     Circle
Helisoma antrosa (Conrad)       Circle     Circle Circle    
Helisoma trivolvis (Say)       Circle Circle   Circle Circle    
Physa anatina Lea       Circle         Circle Circle
Pupilla blandi Morse       Circle Circle          
Pupilla muscorum (Linne)   Circle       Circle   Circle    
Pupoides albilabris (C. B. Adams)         Circle       Circle  
Retinella electrina (Gould) Circle     Circle       Circle    
Sphaerium solidulum (Prime) Circle     Circle Circle          
Strobilops sparsicosta Baker           Circle     Circle Circle
Succinea grosvenori Lea Circle   Circle Circle Circle Circle Circle      
Vallonia gracilicosta Reinhardt   Circle Circle Circle Circle   Circle Circle    
Vallonia pulchella (Muller)       Circle            
Vertigo tridentata Wolf   Circle       Circle     Circle  

Plate 16--A representative assemblage of mollusks from Illinoian deposits in Kansas. All figures enlarged approximately 5 times natural size. An Acrobat PDF version of this plate is available that shows more detail.

  1. Gastrocopta armifera (Say)
  2. Pupilla muscorum (Linne)
  3. Pupilla blandi Morse
  4. Vertigo tridentata Wolf
  5. Gastrocopta tappaniana (C. B. Adams)
  6. Gastrocopta holzingeri (Sterki)
  7. Vallonia gracilicosta Reinhardt
  8. Vallonia pulchella (Müller)
  9. Helicodiscus singleyanus (Pilsbry)
  10. Deroceras laeve (Müller)
  11. Pupoides albilabris (C. B. Adams)
  12. Retinella electrina (Gould)
  13. HeIicodiscus parallelus (Say)
  14. Hawaiia minuscula (Binney)
  15. Strobilops sparsicosta Baker
  16. Carychium exiguum (Say)
  17. Discus cronkhitei (Newcomb)
  18. Helisoma antrosa (Conrad)
  19. Physa anatina Lea
  20. Sphaerium solidulum Prime
  21. Gyraulus similaris Baker
  22. Helisoma trivolvis (Say)
  23. Succinea grosvenori Lea
Black and white plate of samples

Typically, Crete sands and gravels grade upward into fluviatile silts, which in turn grade into massive, well-sorted silts judged to be eolian in origin. Almost everywhere in the State where these sediments can be recognized, as well as in Nebraska, Iowa, Illinois, and other surrounding states, the Crete-Loveland sequence has developed in its upper part the Sangamon soil. The Sangamon soil is in fact the most reliable stratigraphic datum available for the recognition of Crete-Loveland deposits, and the fauna under present consideration has been in every case collected only where the Sangamon soil was recognizable as a stratigraphic reference.

The Illinoian assemblage of mollusks is less distinctive than that in Nebraskan, Kansan, or Wisconsinan deposits primarily because no segment of the assemblage is restricted to it. However, with care and a reasonably complete faunule, an Illinoian assemblage can be distinguished from other aggregations of Pleistocene mollusks in Kansas. In general, the Illinoian assemblage is intermediate in character between that in Kansan sediments stratigraphically below it and that in Wisconsinan deposits stratigraphically above it. But as inferred above, the age of the assemblage is based largely on its relation to the Sangamon soil.

In spite of its intermediate character, the Illinoian faunal assemblage has a number of distinctive features worthy of note.

(1) About 14 species of common occurrence in Kansan deposits do not appear in the Illinoian assemblages. None of the genera of branchiate gastropods, such as Amnicola, Pomatiopsis, and Valvata seems to have survived the Yarmouthian interglacial interval in the midcontinent region. Likewise, aquatic pulmonate snails, such as Aplexa, Ferrissia, Menetus, Planorbula, and Promenetus, most species of Gyraulus, and the larger species of Lymnaea, failed to survive the ecological changes that followed the close of deposition of Sappa silts.

(2) Four species, common in Kansan faunules, make their final appearance in the geologic column in Kansas in Illinoian sediments; these include Carychium perexiguum, Strobilops sparsicosta, Gyraulus similaris, and Helisoma antrosa. The first two species are extinct, Gyraulus similaris survives in relict populations in montane lakes in Colorado, and Helisoma antrosa, although extinct on the Great Plains, is still a commonly occurring snail in the more humid regions of central and eastern United States.

(3) A number of species, among which may be mentioned Columella alticola and Striatura milium which are relatively common in Wisconsinan loess (Tazewellian zone of the Peoria silt), are not found in Illinoian assemblages. Furthermore, the slug Deroceras aenigma, common in Nebraskan and Kansan faunas, is replaced in Illinoian molluscan assemblages by Deroceras laeve.

The small number of localities from which molluscan assemblages of Illinoian age are reported here is no indication of the extent of Illinoian sediments in the State. At most localities where Illinoian sediments can be recognized, the gravels, sands, or silts are nonfossiliferous. The reason for this is not completely understood, but a number of contributing factors are known. It is not surprising that few instances of fossiliferous Crete sand and gravel are known, because the fragile shells of mollusks cannot survive for long the abrasive action of coarse sediments in transit. In fact, it is remarkable that any examples of fossiliferous Crete materials are known, since there is not a single instance known to us of shells occurring in recognizable faunules in the coarse phases of the Holdrege or Grand Island gravels. Another contributing factor is that generally over the State, while Illinoian sediments are widespread, they exist as relatively thin strata. Not infrequently, the weathering that produced the Sangamon soil has progressed completely through the entire Illinoian sequence, which would have destroyed, of course, any shells previously present. Finally, the climate during Illinoian time may have been unfavorable to mollusks, although this seems unlikely in view of the kinds that are known to be present. The absence of molluscan remains in unleached Loveland loess at a number of localities remains unexplained.

It is difficult to escape the conclusion that a profound change in ecological conditions in the Great Plains occurred during or at the close of the Yarmouthian interglacial interval, or at the beginning of the Illinoian cycle of erosion and deposition. Wholesale extinction of great populations of branchiate and other gastropods adapted to life in permanent water, which thrived in western Kansas in late Kansan and early Yarmouthian time, is indicative of a less humid environment, or at least of a marked decline in the prevalence of permanent ponds and lakes of clear water in the Great Plains region. The assemblage of aquatic gastropods in late Kansan silt, the prevalence in them of zygospores of some Chara-like alga, and abundance at many exposures of the valves of ostracods all point toward an environment of permanent slow-flowing or ponded water, without excessive siltation, and perhaps with an average temperature somewhat below that prevalent in the same region today or in Illinoian time. By contrast, the aquatic gastropods in Illinoian deposits indicate an environment of ephemeral ponds and silt-laden streams. Anodonta, a lammellibranch pelecypod, and the pulmonate gastropods Helisoma trivolvis and Physa anatina, as well as the little clam Sphaerium solidulum, are all animals capable of living in heavily silted ponds or streams, and capable also of surviving considerable periods without open water. Gyraulus similaris probably was limited to spring-fed ponds or other cooler and clearer waters.

With the exception of Strobilops sparsicosta, which probably lived in or near woodlands, the terrestrial fauna associated with Illinoian deposits is adapted for life in prairies, meadows, or perhaps woodland borders or in small areas of shrubs. Gastrocopta armifera, Hawaiia minuscula, and Pupoides albilabris live in the prairies of western Kansas today, and survive periods of great aridity. Pupilla muscorum, however, and Retinella electrina, Carychium perexiguum, C. exiguum, Helicodiscus singleyanus, Vallonia gracilicosta, Vertigo tridentata, and others are absent from the living molluscan fauna in the Great Plains, so it must be conceded that in Illinoian time ecological conditions were somewhat more favorable toward mollusks than those prevalent today. Certainly the contrast between ecological conditions in Illinoian time and Kansan as well as Nebraskan time was much greater than that between Illinoian and modern time, and we judge that climatic conditions in Kansas during the Illinoian Age were not significantly different than present-day conditions, except for slightly greater rainfall and greater and more widespread sedimentation. It seems, for example, that the ecological situation in Norton or Phillips County in Illinoian time might have been not unlike that in Dickinson or Marion County today.

Molluscan Faunal Assemblages in Wisconsinan Deposits

The molluscan faunal assemblage associated with Wisconsinan sediments has been previously treated by Frye and A. B. Leonard (1951), and by A. B. Leonard (1951, 1952). In the present report, the Wisconsinan molluscan fauna is divided into pre- and post-Bradyan assemblages; the occurrence and distribution of 28 kinds of mollusks from pre-Bradyan Wisconsinan silts at 137 localities is shown on Figure 9, and illustrations of a representative assemblage appear on Plate 17. The occurrence and distribution of 16 species of mollusks at 27 localities where post-Bradyan Wisconsinan deposits have been studied is shown on Figure 10, while a representative assemblage of these mollusks is illustrated on Plate 18.

Figure 9--Occurrence of fossil mollusks in pre-Bradyan Wisconsinan deposits at 137 localities in Kansas. Asterisk in location indicates collection from hand auger samples. An Acrobat PDF version of this figure is available.

Molluscan species Faunal localities
SW sec. 30, T. 3 S., R. 38 W., Cheyenne SE sec. 28, T. 3 S., R. 39 W., Cheyenne SE sec. 17, T. 1 S., R. 41 W., Cheyenne SE sec. 16, T. 1 S., R. 42 W., Cheyenne NW sec. 6, T. 30 S., R. 23 W., Clark NW sec. 20, T. 33 S., R. 23 W., Clark NE sec. 26, T. 33 S., R. 24 W., Clark NW sec. 31, T. 5 S., R. 3 W., Cloud NE sec. 3, T. 3 S., R. 27 W., Decatur NE sec. 6, T. 3 S., R. 27 W., Decatur NW sec. 35, T. 3 S., R. 27 W., Decatur SW sec. 23, T. 4 S., R. 27 W., Decatur SW sec. 26, T. 4 S., R. 27 W., Decatur SW sec. 7, T. 5 S., R. 27 W., Decatur
Gastrocopta holzingeri (Sterki)
Succinea ovalis Say Circle
Gastrocopta armifera (Say)
Hendersonia occulta (Say) Circle
Discus shimeki (Pilsbry) Circle Circle Circle Circle Circle
Succinea grosvenori Lea Circle Circle Circle Circle Circle Circle Circle Circle
Vertigo modesta (Say) Circle Circle Circle Circle
Discus cronkhitei (Newcomb) Circle Circle Circle Circle
Columella alticola (Ingersoll) Circle
Striatura milium (Morse) Circle
Retinella electrina (Gould)
Cionella lubrica (Müller)
Vertigo gouldi coloradensis (Cockerell) Circle
Zonitoides arboreus (Say)
Molluscan species Faunal localities
NE sec. 1, T. 3 S., R. 28 W., Decatur NW sec. 23, T. 5 S., R. 28 W., Decatur NE sec. 14, T. 5 S., R. 29 W., Decatur SE sec. 6, T. 2 S., R. 20 E., Doniphan NE sec. 8, T. 1 S., R. 19 E., Doniphan SW sec. 29, T. 15 S., R. 18 W., Ellis SW sec. 18, T. 14 S., R. 19 W., Ellis SW sec. 24, T 21 S., R. 29 W., Finney NE sec. 34, T. 22 S., R. 30 W., Finney SE sec. 4, T. 24 S., R. 32 W., Finney SW sec. 9, T. 24 S., R. 32 W., Finney NW sec. 16, T. 29 S., R. 24 W., Ford NE sec. 2, T. 27 S., R. 22 W., Ford NE sec. 35, T. 15 S., R. 27 W., Gove
Gastrocopta holzingeri (Sterki) Circle Circle
Succinea ovalis Say Circle Circle
Gastrocopta armifera (Say) Circle
Hendersonia occulta (Say)
Discus shimeki (Pilsbry) Circle Circle Circle Circle Circle
Succinea grosvenori Lea Circle Circle Circle Circle
Vertigo modesta (Say) Circle Circle
Discus cronkhitei (Newcomb) Circle Circle Circle Circle
Columella alticola (Ingersoll) Circle
Striatura milium (Morse) Circle
Retinella electrina (Gould) Circle
Cionella lubrica (Müller) Circle
Vertigo gouldi coloradensis (Cockerell) Circle
Zonitoides arboreus (Say) Circle Circle Circle
Molluscan species Faunal localities
SW sec. 36, T. 14 S., R. 29 W., Gove SW sec. 23, T. 9 S., R. 21 W., Graham NE sec. 28, T. 9 S., R. 22 W., Graham SE sec. 28, T. 27 S., R. 35 W., Grant *NE sec. 1, T. 25 S., R. 29 W., Gray NW sec. 9, T. 16 S., R. 40 W., Greeley *NW sec. 19, T. 21 S., R. 40 W., Hamilton SE sec. 12, T. 25 S., R. 41 W., Hamilton SW sec. 36, T. 25 S., R. 41 W., Hamilton SE sec. 35, T. 35 S., R. 8 W., Harper NW sec. 3, T. 28 S., R. 32 W., Haskell NE sec. 22, T. 30 S., R. 34 W., Haskell NE sec. 27, T. 23 S., R. 22 W., Hodgeman NW sec. 26, T. 23 S., R. 25 W., Hodgeman
Gastrocopta holzingeri (Sterki)
Succinea ovalis Say
Gastrocopta armifera (Say) Circle
Hendersonia occulta (Say)
Discus shimeki (Pilsbry) Circle
Succinea grosvenori Lea Circle Circle
Vertigo modesta (Say)
Discus cronkhitei (Newcomb) Circle
Columella alticola (Ingersoll)
Striatura milium (Morse)
Retinella electrina (Gould)
Cionella lubrica (Müller)
Vertigo gouldi coloradensis (Cockerell) Circle
Zonitoides arboreus (Say)
Molluscan species Faunal localities
NE sec. 4, T. 1 S., R. 7 W., Jewell SW sec. 19, T. 5 S., R. 8 W., Jewell *NW sec. 31, T. 23 S., R. 38 W., Kearny NW sec. 3, T. 28 S., R. 9 W., Kingman SW sec. 18, T. 16 S., R. 28 W., Lane NW sec. 7, T. 20 S., R. 28 W., Lane SE sec. 29, T. 20 S., R. 28 W., Lane SW sec. 2, T. 16 S., R. 29 W., Lane NW sec. 3, T. 16 S., R. 29 W., Lane NE sec. 15, T. 16 S., R. 29 W., Lane SE sec. 15, T. 16 S., R. 30 W, Lane *NE sec. 17, T. 17 S., R. 30 W., Lane SE sec. 27, T. 11 S., R. 32 W., Logan NW sec. 13, T. 12 S., R. 32 W., Logan
Gastrocopta holzingeri (Sterki)
Succinea ovalis Say
Gastrocopta armifera (Say) Circle
Hendersonia occulta (Say) Circle Circle Circle
Discus shimeki (Pilsbry) Circle Circle
Succinea grosvenori Lea Circle Circle Circle Circle Circle
Vertigo modesta (Say) Circle
Discus cronkhitei (Newcomb) Circle Circle
Columella alticola (Ingersoll) Circle
Striatura milium (Morse) Circle
Retinella electrina (Gould) Circle Circle
Cionella lubrica (Müller) Circle Circle
Vertigo gouldi coloradensis (Cockerell)
Zonitoides arboreus (Say)
Molluscan species Faunal localities
SE sec. 3, T. 14 S., R. 32 W., Logan NW sec. 27, T. 15 S., R. 32 W., Logan NE sec. 2, T. 14 S., R. 35 W., Logan NE sec. 9, T. 15 S., R. 35 W., Logan SE sec. 3, T. 11 S., R. 26 W., Logan NE sec. 32, T. 12 S., R. 37 W., Logan NE sec. 8, T. 33 S., R. 27 W., Meade NE sec. 11, T. 30 S., R. 28 W., Meade SW sec. 30, T. 31 S., R. 28 W., Meade NW sec. 6, T. 30 S., R. 30 W., Meade NW sec. 28, T. 8 S., R. 8 W., Mitchell NW sec. 10, T. 16 S., R. 22 W., Ness NE sec. 16, T. 16 S., R. 22 W., Ness NE sec. 29, T. 16 S., R. 24 W., Ness
Gastrocopta holzingeri (Sterki)
Succinea ovalis Say
Gastrocopta armifera (Say) Circle
Hendersonia occulta (Say) Circle
Discus shimeki (Pilsbry) Circle
Succinea grosvenori Lea
Vertigo modesta (Say) Circle
Discus cronkhitei (Newcomb)
Columella alticola (Ingersoll)
Striatura milium (Morse)
Retinella electrina (Gould)
Cionella lubrica (Müller)
Vertigo gouldi coloradensis (Cockerell)
Zonitoides arboreus (Say)
Molluscan species Faunal localities
NW sec. 26, T. 2 S., R. 23 W., Norton NW sec. 11, T. 3 S., R. 23 W., Norton NE sec. 10, T. 4 S., R. 23 W., Norton SE sec. 27, T. 4 S., R. 23 W., Norton SE sec. 13, T. 1 S., R. 24 W., Norton NE sec. 7, T. 5 S., R. 24 W., Norton SE sec. 5, T. 2 S., R. 17 W., Phillips NW sec. 23, T. 2 S., R. 18 W., Phillips NE sec. 25, T. 1 S., R. 20 W., Phillips NE sec. 2, T. 3 S., R. 31 W., Rawlins NW sec. 8, T. 5 S., R. 31 W., Rawlins SW sec. 2, T. 3 S., R. 33 W.,, Rawlins NE sec. 9, T. 3 S., R. 33 W., Rawlins NW sec. 20, T. 4 S., R. 33 W., Rawlins
Gastrocopta holzingeri (Sterki)
Succinea ovalis Say
Gastrocopta armifera (Say) Circle
Hendersonia occulta (Say)
Discus shimeki (Pilsbry) Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Succinea grosvenori Lea Circle Circle Circle Circle Circle Circle Circle Circle
Vertigo modesta (Say) Circle Circle Circle Circle Circle Circle Circle
Discus cronkhitei (Newcomb) Circle Circle Circle Circle Circle Circle
Columella alticola (Ingersoll) Circle Circle
Striatura milium (Morse) Circle Circle Circle
Retinella electrina (Gould) Circle Circle
Cionella lubrica (Müller) Circle
Vertigo gouldi coloradensis (Cockerell)
Zonitoides arboreus (Say) Circle Circle Circle
Molluscan species Faunal localities
NE sec. 8, T. 3 S., R. 34 W., Rawlins NE sec. 29, T. 3 S., R. 36 W., Rawlins NW sec. 5, T. 3 S., R. 4 W., Republic SE sec. 7, T. 10 S., R. 16 W., Rooks SW sec. 30, T. 9 S., R. 19 W., Rooks SW sec. 28, T. 17 S., R. 19 W, Rush SE sec. 17, T. 18 S., R. 31 W., Scott SE sec. 6, T. 17 S., R. 32 W., Scott *SW sec. 31, T. 20 S., R. 34 W., Scott SW sec. 15, T. 6 S., R. 28 W., Sheridan SW sec. 3, T. 7 S., R. 28 W., Sheridan SE sec. 28, T. 7 S., R. 28 W., Sheridan NW sec. 34, T. 8 S., R. 28 W., Sheridan SW sec. 10, T. 9 S., R. 28 W., Sheridan
Gastrocopta holzingeri (Sterki)
Succinea ovalis Say
Gastrocopta armifera (Say) Circle Circle Circle
Hendersonia occulta (Say) Circle
Discus shimeki (Pilsbry) Circle Circle Circle Circle Circle Circle
Succinea grosvenori Lea Circle Circle Circle Circle Circle Circle Circle
Vertigo modesta (Say) Circle Circle Circle
Discus cronkhitei (Newcomb) Circle Circle Circle Circle
Columella alticola (Ingersoll)
Striatura milium (Morse) Circle Circle
Retinella electrina (Gould) Circle Circle
Cionella lubrica (Müller) Circle
Vertigo gouldi coloradensis (Cockerell) Circle
Zonitoides arboreus (Say) Circle Circle
Molluscan species Faunal localities
NE sec. 33, T. 10 S., R. 28 W., Sheridan NW sec. 19, T. 8 S., R. 29 W., Sheridan NE sec. 13, T. 6 S., R. 30 W., Sheridan NW sec. 14, T. 6 S., R. 30 W., Sheridan NW sec. 15, T. 6 S., R. 30 W., Sheridan NW sec. 31, T. 8 S., R. 30 W., Sheridan SE sec. 34, T. 10 S., R. 37 W., Sherman SW sec. 26, T. 10 S., R. 38 W., Sherman NW sec. 32, T. 7 S., R. 39 W., Sherman SW sec. 36, T. 6 S., R. 39 W., Sherman SW sec. 20, T. 7 S., R. 40 W., Sherman SE sec. 28, T. 1 S., R. 13 W., Smith SW sec. 11, T. 27 S., R. 39 W., Stanton NE sec. 2, T. 29 S., R. 40 W., Stanton
Gastrocopta holzingeri (Sterki) Circle
Succinea ovalis Say
Gastrocopta armifera (Say) Circle
Hendersonia occulta (Say)
Discus shimeki (Pilsbry) Circle
Succinea grosvenori Lea Circle Circle Circle Circle Circle
Vertigo modesta (Say) Circle
Discus cronkhitei (Newcomb) Circle Circle Circle Circle
Columella alticola (Ingersoll) Circle
Striatura milium (Morse) Circle
Retinella electrina (Gould) Circle
Cionella lubrica (Müller) Circle
Vertigo gouldi coloradensis (Cockerell)
Zonitoides arboreus (Say) Circle
Molluscan species Faunal localities
NW sec. 12, T. 28 S., R. 41 W., Stanton SE sec. 25, T. 29 S., R. 41 W., Stanton SE sec. 21, T. 7 S., R. 31 W., Thomas NE sec. 27, T. 7 S., R. 31 W., Thomas NW sec. 10, T. 8 S., R. 32 W., Thomas NE sec. 17, T. 8 S., R. 32 W., Thomas NW sec. 31, T. 7 S., R. 33 W., Thomas NW sec. 18, T. 9 S., R. 33 W., Thomas SE sec. 27, T. 10 S., R. 33 W., Thomas NW sec. 8, T. 8 S., R. 35 W., Thomas SW sec. 8, T. 8 S., R. 35 W., Thomas SW sec. 32, T. 7 S., R. 36 W., Thomas SW sec. 7, T. 8 S., R. 36 W., Thomas SW sec. 19, T. 8 S., R. 36 W., Thomas
Gastrocopta holzingeri (Sterki)
Succinea ovalis Say
Gastrocopta armifera (Say) Circle
Hendersonia occulta (Say)
Discus shimeki (Pilsbry) Circle Circle
Succinea grosvenori Lea Circle Circle Circle
Vertigo modesta (Say) Circle Circle Circle
Discus cronkhitei (Newcomb) Circle Circle Circle Circle
Columella alticola (Ingersoll) Circle Circle
Striatura milium (Morse)
Retinella electrina (Gould) Circle
Cionella lubrica (Müller)
Vertigo gouldi coloradensis (Cockerell) Circle
Zonitoides arboreus (Say) Circle
Molluscan species Faunal localities
NW sec. 3, T. 11 S., R. 23 W., Trego SW sec. 5, T. 12 S., R. 24 W., Trego NW sec. 26, T. 15 S., R. 38 W., Wallace SW sec. 35, T. 11 S., R. 40 W., Wallace SW sec. 23, T. 12 S., R. 40 W., Wallace NW sec. 1, T. 15 S., R. 40 W., Wallace *SE sec. 35, T. 20 S., R. 35 W., Wichita SW sec. 12, T. 17 S., R. 37 W., Wichita SE sec. 1, T. 19 S., R. 37 W., Wichita NE sec. 12, T. 19 S., R. 37 W., Wichita SE sec. 32, T. 16 S., R. 38 W., Wichita
Gastrocopta holzingeri (Sterki)
Succinea ovalis Say
Gastrocopta armifera (Say)
Hendersonia occulta (Say)
Discus shimeki (Pilsbry) Circle
Succinea grosvenori Lea Circle
Vertigo modesta (Say) Circle
Discus cronkhitei (Newcomb)
Columella alticola (Ingersoll)
Striatura milium (Morse)
Retinella electrina (Gould)
Cionella lubrica (Müller)
Vertigo gouldi coloradensis (Cockerell)
Zonitoides arboreus (Say)

Molluscan species Faunal localities
SW sec. 30, T. 3 S., R. 38 W., Cheyenne SE sec. 28, T. 3 S., R. 39 W., Cheyenne SE sec. 17, T. 1 S., R. 41 W., Cheyenne SE sec. 16, T. 1 S., R. 42 W., Cheyenne NW sec. 6, T. 30 S., R. 23 W., Clark NW sec. 20, T. 33 S., R. 23 W., Clark NE sec. 26, T. 33 S., R. 24 W., Clark NW sec. 31, T. 5 S., R. 3 W., Cloud NE sec. 3, T. 3 S., R. 27 W., Decatur NE sec. 6, T. 3 S., R. 27 W., Decatur NW sec. 35, T. 3 S., R. 27 W., Decatur SW sec. 23, T. 4 S., R. 27 W., Decatur SW sec. 26, T. 4 S., R. 27 W., Decatur SW sec. 7, T. 5 S., R. 27 W., Decatur
Vertigo milium (Gould) Circle Circle
Vertigo tridentata Wolf Circle
Carychium exiguum (Say) Circle
Deroceras laeve (Müller) Circle
Helicodiscus singleyanus (Pilsbry) Circle
Vertigo gouldi paradoxa Sterki Circle Circle Circle Circle
Helicodiscus parallelus (Say) Circle Circle Circle Circle
Euconulus fulvus (Müller) Circle Circle Circle Circle Circle
Hawaiia minuscula (Binney) Circle Circle Circle Circle Circle Circle Circle
Pupilla muscorum (Linne) Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Pupilla blandi Morse Circle Circle Circle Circle Circle Circle
Vallonia gracilicosta Reinhardt Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Succinea avara Say Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Lymnaea parva Lea Circle Circle
Molluscan species Faunal localities
NE sec. 1, T. 3 S., R. 28 W., Decatur NW sec. 23, T. 5 S., R. 28 W., Decatur NE sec. 14, T. 5 S., R. 29 W., Decatur SE sec. 6, T. 2 S., R. 20 E., Doniphan NE sec. 8, T. 1 S., R. 19 E., Doniphan SW sec. 29, T. 15 S., R. 18 W., Ellis SW sec. 18, T. 14 S., R. 19 W., Ellis SW sec. 24, T 21 S., R. 29 W., Finney NE sec. 34, T. 22 S., R. 30 W., Finney SE sec. 4, T. 24 S., R. 32 W., Finney SW sec. 9, T. 24 S., R. 32 W., Finney NW sec. 16, T. 29 S., R. 24 W., Ford NE sec. 2, T. 27 S., R. 22 W., Ford NE sec. 35, T. 15 S., R. 27 W., Gove
Vertigo milium (Gould) Circle Circle Circle Circle Circle
Vertigo tridentata Wolf Circle Circle
Carychium exiguum (Say)
Deroceras laeve (Müller) Circle Circle
Helicodiscus singleyanus (Pilsbry) Circle Circle
Vertigo gouldi paradoxa Sterki Circle Circle
Helicodiscus parallelus (Say) Circle Circle
Euconulus fulvus (Müller) Circle Circle Circle Circle Circle
Hawaiia minuscula (Binney) Circle Circle Circle Circle Circle Circle Circle Circle Circle
Pupilla muscorum (Linne) Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Pupilla blandi Morse Circle Circle Circle Circle Circle Circle Circle Circle Circle
Vallonia gracilicosta Reinhardt Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Succinea avara Say Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Lymnaea parva Lea Circle Circle
Molluscan species Faunal localities
SW sec. 36, T. 14 S., R. 29 W., Gove SW sec. 23, T. 9 S., R. 21 W., Graham NE sec. 28, T. 9 S., R. 22 W., Graham SE sec. 28, T. 27 S., R. 35 W., Grant *NE sec. 1, T. 25 S., R. 29 W., Gray NW sec. 9, T. 16 S., R. 40 W., Greeley *NW sec. 19, T. 21 S., R. 40 W., Hamilton SE sec. 12, T. 25 S., R. 41 W., Hamilton SW sec. 36, T. 25 S., R. 41 W., Hamilton SE sec. 35, T. 35 S., R. 8 W., Harper NW sec. 3, T. 28 S., R. 32 W., Haskell NE sec. 22, T. 30 S., R. 34 W., Haskell NE sec. 27, T. 23 S., R. 22 W., Hodgeman NW sec. 26, T. 23 S., R. 25 W., Hodgeman
Vertigo milium (Gould) Circle
Vertigo tridentata Wolf
Carychium exiguum (Say)
Deroceras laeve (Müller)
Helicodiscus singleyanus (Pilsbry) Circle Circle Circle
Vertigo gouldi paradoxa Sterki Circle Circle Circle Circle
Helicodiscus parallelus (Say) Circle Circle Circle
Euconulus fulvus (Müller) Circle
Hawaiia minuscula (Binney) Circle Circle Circle Circle Circle Circle Circle Circle
Pupilla muscorum (Linne) Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Pupilla blandi Morse Circle Circle Circle Circle Circle Circle Circle Circle Circle
Vallonia gracilicosta Reinhardt Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Succinea avara Say Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Lymnaea parva Lea Circle
Molluscan species Faunal localities
NE sec. 4, T. 1 S., R. 7 W., Jewell SW sec. 19, T. 5 S., R. 8 W., Jewell *NW sec. 31, T. 23 S., R. 38 W., Kearny NW sec. 3, T. 28 S., R. 9 W., Kingman SW sec. 18, T. 16 S., R. 28 W., Lane NW sec. 7, T. 20 S., R. 28 W., Lane SE sec. 29, T. 20 S., R. 28 W., Lane SW sec. 2, T. 16 S., R. 29 W., Lane NW sec. 3, T. 16 S., R. 29 W., Lane NE sec. 15, T. 16 S., R. 29 W., Lane SE sec. 15, T. 16 S., R. 30 W, Lane *NE sec. 17, T. 17 S., R. 30 W., Lane SE sec. 27, T. 11 S., R. 32 W., Logan NW sec. 13, T. 12 S., R. 32 W., Logan
Vertigo milium (Gould) Circle Circle Circle Circle
Vertigo tridentata Wolf
Carychium exiguum (Say) Circle Circle
Deroceras laeve (Müller)
Helicodiscus singleyanus (Pilsbry)
Vertigo gouldi paradoxa Sterki Circle
Helicodiscus parallelus (Say) Circle
Euconulus fulvus (Müller) Circle Circle
Hawaiia minuscula (Binney) Circle Circle Circle Circle Circle Circle Circle Circle Circle
Pupilla muscorum (Linne) Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Pupilla blandi Morse Circle Circle Circle Circle Circle Circle Circle
Vallonia gracilicosta Reinhardt Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Succinea avara Say Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Lymnaea parva Lea Circle Circle Circle
Molluscan species Faunal localities
SE sec. 3, T. 14 S., R. 32 W., Logan NW sec. 27, T. 15 S., R. 32 W., Logan NE sec. 2, T. 14 S., R. 35 W., Logan NE sec. 9, T. 15 S., R. 35 W., Logan SE sec. 3, T. 11 S., R. 26 W., Logan NE sec. 32, T. 12 S., R. 37 W., Logan NE sec. 8, T. 33 S., R. 27 W., Meade NE sec. 11, T. 30 S., R. 28 W., Meade SW sec. 30, T. 31 S., R. 28 W., Meade NW sec. 6, T. 30 S., R. 30 W., Meade NW sec. 28, T. 8 S., R. 8 W., Mitchell NW sec. 10, T. 16 S., R. 22 W., Ness NE sec. 16, T. 16 S., R. 22 W., Ness NE sec. 29, T. 16 S., R. 24 W., Ness
Vertigo milium (Gould) Circle
Vertigo tridentata Wolf
Carychium exiguum (Say)
Deroceras laeve (Müller)
Helicodiscus singleyanus (Pilsbry) Circle Circle
Vertigo gouldi paradoxa Sterki Circle Circle Circle
Helicodiscus parallelus (Say) Circle
Euconulus fulvus (Müller) Circle
Hawaiia minuscula (Binney) Circle Circle Circle Circle Circle Circle Circle Circle
Pupilla muscorum (Linne) Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Pupilla blandi Morse Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Vallonia gracilicosta Reinhardt Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Succinea avara Say Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Lymnaea parva Lea Circle Circle
Molluscan species Faunal localities
NW sec. 26, T. 2 S., R. 23 W., Norton NW sec. 11, T. 3 S., R. 23 W., Norton NE sec. 10, T. 4 S., R. 23 W., Norton SE sec. 27, T. 4 S., R. 23 W., Norton SE sec. 13, T. 1 S., R. 24 W., Norton NE sec. 7, T. 5 S., R. 24 W., Norton SE sec. 5, T. 2 S., R. 17 W., Phillips NW sec. 23, T. 2 S., R. 18 W., Phillips NE sec. 25, T. 1 S., R. 20 W., Phillips NE sec. 2, T. 3 S., R. 31 W., Rawlins NW sec. 8, T. 5 S., R. 31 W., Rawlins SW sec. 2, T. 3 S., R. 33 W.,, Rawlins NE sec. 9, T. 3 S., R. 33 W., Rawlins NW sec. 20, T. 4 S., R. 33 W., Rawlins
Vertigo milium (Gould) Circle
Vertigo tridentata Wolf
Carychium exiguum (Say)
Deroceras laeve (Müller) Circle
Helicodiscus singleyanus (Pilsbry) Circle Circle Circle
Vertigo gouldi paradoxa Sterki Circle Circle Circle
Helicodiscus parallelus (Say) Circle Circle Circle Circle Circle Circle
Euconulus fulvus (Müller) Circle Circle Circle Circle Circle Circle Circle
Hawaiia minuscula (Binney) Circle Circle Circle Circle Circle Circle Circle Circle
Pupilla muscorum (Linne) Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle Circle
Pupilla blandi Morse Circle Circle Circle Circle Circle